In the ever-evolving world of Teyvat, the Sprayfeather Gill emerges as a fascinating coral, not merely a feather-shaped seagrass, exclusively found in Natlan's aquatic realms. But why should Travelers prioritize collecting this unique resource in 2025? It serves as a vital Ascension material for characters like Mualani, essential for maxing out their levels and enhancing team compositions. With a respawn timer of 48 hours, similar to other local specialties, efficient farming becomes crucial to avoid unnecessary delays. This guide provides a comprehensive walkthrough based on the latest in-game data, ensuring players can navigate the waters of Toyac Springs with precision and ease. After all, isn't it better to spend time battling formidable foes rather than endlessly searching for resources?

Total Locations and Key Areas

Sprayfeather Gills are abundant yet strategically clustered, with a total of 75 scattered across Toyac Springs. They primarily grow in groups of two or three, making group collection efficient. To access all areas, players must first complete specific quests; for instance, the "Tale of Dreams Plucked From Fire" Quest is mandatory for the Underground Lava River locations. Here's a quick overview of the distribution:

Area	Number of Gills	Access Requirements
Ameyalco Waters	48	None (freely accessible)
The Upper Sanctum	12	None
Underground Lava River: Lower Level	15	Complete "Tale of Dreams Plucked From Fire" Quest

This table highlights the hotspots, but remember, why struggle through blind searches when a systematic approach can save hours? Now, let's dive into the step-by-step collection methods for each area.

Detailed Collection Guide for Ameyalco Waters

Ameyalco Waters holds the majority of Sprayfeather Gills, demanding careful navigation. Players should start by fast-traveling to key waypoints and transform into the Koholasauri for aquatic mobility—this form is indispensable for swift swimming. Below are the optimized routes:

• Northern Section:

• Teleport to Toyac Springs' northern waypoint.

• Head east toward the water to collect three Sprayfeather Gills.

• Continue moving east to pick up another one.

• Transform into Koholasauri, swim south, and gather three more.

• Move west to acquire two additional Gills.

• Proceed to western small ponds to find seven scattered pieces. Isn't it satisfying to clear a cluster effortlessly?

• Central and Eastern Sections:

• Fast-travel to Ameyalco Waters' northern Waypoint.

• Glide west to collect three Sprayfeather Gills.

• Move south for another two.

• As Koholasauri, swim to the northeastern shore for three Gills.

• Head southwest to pick four more.

• Swim to the western small island for three Gills, then further west for one additional.

• Finally, move toward the southern shore for two, and swim east for the last one in this segment.

• Southwestern and Island Sections:

• Teleport to Ameyalco Waters' southwestern island.

• Transform into Koholasauri and head to the northwestern shore for three Gills.

• Go northeast to collect four.

• Move east in front of the Shrine of Depths for four more—imagine the rewards after such a trek! 👍

• Swim south then east to acquire six Sprayfeather Gills.

• Approach the small southern island for two Gills.

• Swim to the tiny northeastern island for the final two, completing the Ameyalco Waters haul.

Collection in The Upper Sanctum

The Upper Sanctum offers fewer Gills but requires aerial maneuvers. Start by teleporting to the western Waypoint:

• Collect two Sprayfeather Gills immediately.

• Walk east to the end of the flying land, then glide toward the eastern land.

• Upon reaching ponds, gather three more Gills. 🔄

• Fast-travel to the eastern Waypoint and pick three from the adjacent pond.

• Walk to the end of the flying land, leap northwest, and secure four Gills beside the water. Simple, right?

Underground Lava River: Lower Level Guide

This area is locked behind quests but rewards persistent players. Begin by teleporting to the Waypoint unlocked during the "Tracer No Tracing" Quest:

• Transform into Koholasauri and swim south in the lava.

• Use the Jet-Set Eddie (vortex) to reach the floating Monetoo.

• Guide Monetoo back to its location and infuse with Phlogiston.

• Proceed behind the unlocked door to collect five Sprayfeather Gills. How rewarding is it to overcome such challenges?

Pro Tips for Efficient Farming

To maximize yield, consider these strategies based on current meta in 2025:

• Character Recommendations: Use ranged characters like Fischl for quick elemental interactions, as suggested in general guides. Mualani benefits directly, so prioritize her ascension once Gills are farmed. 😊

• Respawn Management: Since Gills respawn every 48 hours, mark collection times in-game. Plan routes during cooldowns to cover all areas—e.g., start with Ameyalco Waters, then move to others.

• General Best Practices:

• Always transform into Koholasauri for faster swimming.

• Equip artifacts that boost movement speed in water.

• Farm in co-op mode to share resources, but ensure quest prerequisites are met solo.

In conclusion, mastering Sprayfeather Gill collection in 2025 transforms a tedious task into a streamlined adventure. With 75 Gills spread across Toyac Springs, focusing on clustered groups and leveraging transformations ensures players can swiftly ascend characters like Mualani. Remember, efficiency isn't just about speed—it's about reclaiming time for epic battles and new quests in Genshin Impact's ever-expanding world. 🔥